The swallows and martins are a group of passerine birds in the family Hirundinidae which are characterised by their adaptation to aerial feeding. Swallow is used colloquially in Europe as a synonym for the Barn Swallow.
This family comprises two subfamilies: Pseudochelidoninae (the river martins of the genus Pseudochelidon) and Hirundinidae (all other swallows and martins). Within the Hirundinidae, the name "martin" tends to be used for the squarer-tailed species, and the name "swallow" for the more fork-tailed species; however, there is no scientific distinction between these two groups. The family contains around 83 species in 19 genera.
The swallows have a cosmopolitan distribution across the world and breed on all the continents except Antarctica. It is believed that this family originated in Africa as hole-nesters; Africa still has the greatest diversity of species. They also occur on a number of oceanic islands. A number of European and North American species are long-distance migrants; by contrast, the West and South African swallows are non-migratory. A few species of swallow and martin are threatened with extinction by human activities, although other species have benefited from human changes to the environment and live around humans.
The swallows and martins have an evolutionary conservative body shape which is similar across the family but is unlike that of other passerines. Swallows have adapted to hunting insects on the wing by developing a slender streamlined body and long pointed wings, which allow great maneuverability and endurance, as well as frequent periods of gliding. Their body shape allows for very efficient flight, which costs 50-75% less for swallows than equivalent passerines of the same size. Swallows usually forage at around 30–40 km/h, although they are capable of reaching speeds of between 50–65 km/h when traveling.
The bill of the Sand Martin is typical for the family, being short and wide.
Like the unrelated swifts and nightjars, which hunt in a similar way, they have short bills, but strong jaws and a wide gape. Their body length ranges from about 10–24 cm (3.9–9.4 in) and their weight from about 10–60 g (0.35–2.1 oz). The wings are long, pointed, and have nine primary feathers. The tail has 12 feathers and may be deeply forked, somewhat indented, or square-ended. A long tail increases maneuverability, and may also function as a sexual adornment, since the tail is frequently longer in males. In Barn Swallows the tail of the male is 18% longer than the females, and females will select mates on the basis of tail length.
The legs are short, and their feet are adapted for perching rather than walking, as the front toes are partially joined at the base. Swallows are capable of walking and even running, but they do so with a shuffling, waddling gait. The leg muscles of the river martins (Pseudochelidon) are stronger and more robust than those of other swallows.
The most common hirundine plumage is glossy dark blue or green above and plain or streaked underparts, often white or rufous. Species which burrow or live in dry or mountainous areas are often matte brown above (e.g. Sand Martin and Crag Martin). The sexes show limited or no sexual dimorphism, with longer outer tail feathers in the adult male probably being the most common distinction.
For the most part swallows are insectivorous, taking flying insects on the wing. Across the whole family a wide range of insects are taken from most insect groups, but the composition of any one prey type in the diet varies by species and with the time of year. Individual species may be selective, they do not scoop up every insect around them, but instead select larger prey items than would be expected by random sampling. In addition the ease of capture of different insect types affects their rate of predation by swallows. They also avoid certain prey types; in particular stinging insects such as bees and wasps are generally avoided. In addition to insect prey a number of species will occasionally consume fruits and other plant matter. Species in Africa have been recorded eating the seeds of Acacia trees, and these are even fed to the young of the Greater Striped Swallow.
The swallows generally forage for prey that is on the wing, but they will on occasion snap prey off branches or on the ground. The flight may be fast and involve a rapid succession of turns and banks when actively chasing fast moving prey; less agile prey may be caught with a slower more leisurely flight that includes flying in circles and bursts of flapping mixed with gliding. Where several species of swallow feed together they will be separated into different niches based on height off the ground, some species feeding closer to the ground where as other feeding at higher levels. Similar separation occurs where feeding overlaps with swifts. Niche separation may also occur with the size of prey chosen.
The more primitive species nest in existing cavities, for example in an old woodpecker nest, while other species excavate burrows in soft substrate such as sand banks. Swallows in the genera Hirundo, Ptyonoproggne, Cecropis, Petrochelidon and Delichon build mud nests close to overhead shelter in locations that are protected from both the weather and predators. The mud-nesters are most common in the Old World, particularly Africa, whereas cavity-nesters are the rule in the New World. Mud nesting species in particular are limited in areas of high humidity, which causes the mud nests to crumble. Many cave, bank and cliff dwelling species of swallow nest in large colonies. Mud nests are constructed by both males and females, and amongst the tunnel diggers the excavation duties are shared as well. In historical times, the introduction of man-made stone structures such as barns and bridges, together with forest clearance, has led to an abundance of colony sites around the globe, significantly increasing the breeding ranges of some species. Birds living in large colonies typically have to contend with both ectoparasites and conspecific nest parasitism. Old males benefit most from coloniality, since they are able to maintain their own nests and benefit from frequent extra-pair copulations.
Pairs of mated swallows are monogamous, and pairs of non-migratory species often stay near their breeding area all year, though the nest site is defended most vigorously during the breeding season. Migratory species often return to the same breeding area each year, and may select the same nest site if they were previously successful in that location. First-year breeders generally select a nesting site close to where they were born and raised. The breeding of temperate species is seasonal, whereas that of subtropical or tropical species can either be continuous throughout the year or seasonal. Seasonal species in the subtropics or tropics are usually timed to coincide with the peaks in insect activity, which is usually the wet season, but some species like the White-throated Blue Swallow nest in the dry season to avoid flooding in their riverbank nesting habitat. All swallows will defend their nests from egg predators, although solitary species are more aggressive towards predators than colonial species. Overall the contribution of male swallows towards parental care is the highest of any passerine bird.
The eggs of swallows tend to be white, although those of some mud-nesters are speckled. The average clutch size is around four to five eggs in temperate areas and two to three eggs in the tropics. The incubation duties are shared in some species, in others the eggs are incubated solely by the females. Amongst the species where the male helps with incubation the contribution varies amongst species, with some species like the Cliff Swallow sharing the duties equally and the female doing most of the work in others. Amongst the Barn Swallows the male of the American subspecies helps (to a small extent) whereas the European subspecies does not. Even in species where the male does not incubate the eggs the male may sit on them when the female is away to reduce heat loss. Incubation stints last for 5–15 minutes and are followed by bursts of feeding activity. From laying, swallow eggs take between 10–21 days to hatch, with 14–18 days being more typical.
The chicks of swallows hatch naked, generally with only a few tufts of down. The eyes are closed when and do not fully open for up to 10 days. The feathers take a few days to begin to sprout, and the chicks are brooded by the parents until they are able to thermoregulate. On the whole they develop slowly compared to other passerine birds. The parents do not usually feed the chicks individual insects but instead a bolus of food comprising ten to a hundred insects. Regardless of whether the species has males that incubate or brood the chicks the males of all swallows and martins will help feed the chicks. It is difficult to judge when swallows and martins fledge, as they will be enticed out of the nest after three weeks by parents but frequently return to the nest afterwards in order to roost.
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